蕨类植物作为生物进化的重要分支，在生态系统中发挥着不可替代的作用。研究团队以传统分类学为基础，结合分子系统学、生物地理学和细胞学等多学科方法，围绕部分关键类群及生物多样性热点地区，开展系统的蕨类植物多样性研究。

研究团队已前往我国多数省份、以及越南、泰国、老挝、尼泊尔等国开展了大量的野外考察，采集标本万余号，同时收集了对应的数字照片和实验材料。完了高黎贡山、金平分水岭国家自然保护区、河口县、大力苍山和跨境西隆山的蕨类植物名录编写。

截至目前，研究团队与国内外同行合作共同发表了学术论文79篇，其中SCI收录论文77篇；团队负责人作为副主编参与编写学术专著4部。研究成果包括构建了瓶尔小草科（Ophioglossaceae）、凤尾蕨亚科（Pteridoideae）、瓦韦族（Lepisoreae）、薄唇蕨属（Leptochilus）、叉蕨属（Tectaria）等类群的全球系统发育框架；首次结合系统发育和形态学等证据，提出了凤尾蕨属（Pteris）与叉蕨属新的属下分类系统。发表蕨类植物2个新科、11个新属、47个新种及50个新组合。

   

071001-植物学

植物系统分类，蕨类植物多样性，系统进化

2022-08--2024-08   密苏里植物园   访问学者
2010-09--2013-07   中国科学院成都生物研究所   博士
2007-09--2010-07   四川农业大学   硕士
2003-09--2007-07   四川农业大学   学士

1. Zhang, L., Liang, Z.-L., Lu, N.T., Zhou, X.-M., Knapp, R., Pollawatn, R., Zhang, L.-L., Cicuzza, D., Li-Bing Zhang* (2026) Himalayan or tropical Asian origin: new insights into the phylogeny, biogeography, and systematics of the fern genus Selliguea and its allied genera (Polypodiaceae). Journal of Systematics and Evolution. (accepted)
2. Zhang, L., Liang, Z.-L., Fan, X.-P., Lu, N.T., Zhou, X.-M., Wei, H.-J., & Zhang, L.-B.* (2024) The Indo-Burma Biodiversity Hotspot for ferns: Updated phylogeny, hidden diversity, and biogeography of the Java fern genus Leptochilus (Polypodiaceae). Plant Diversity 46: 698–712. https://doi.org/10.1016/j.pld.2024.08.005
3. Zhang, L., Lu, N.T., Zhou, X.-M., Zhou, Z., Thepkayson, K., Luong, T.T., & Zhang, L.-B. (2024) Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot. Phytotaxa 659: 213–35.  https://doi.org/10.11646/phytotaxa.659.3.1
4.  Nie, L.-Y., Zhang, L., Liang, Z.-L., Pollawatn, R., Yan, Y.-H., Lu, N.T., Knapp, R., Wan, X., Cicuzza, D., Chen, X.-X., Chen, H.-F., Wang, A.-H., Liao, Y.-J., Wang, F.-G., & Zhang, L.-B. (2023). Phylogeny, character evolution, and biogeography of the fern genus Bolbitis (Dryopteridaceae). Molecular Phylogenetics and Evolution 178: 107633. https://doi.org/10.1016/j.ympev.2022.107633
5. Zhang, L., & Zhang, L.-B. (2022). Phylogeny, character evolution, and systematics of the fern family Ophioglossaceae based on Sanger sequence data, plastomes, and morphology. Molecular Phylogenetics and Evolution: 107512. https://doi.org/10.1016/j.ympev.2022.107512
6.  Zhang, L., Fan, X.-P., Petchsri, S., Zhou, L., Pollawatn, R., Zhang, X., Zhou, X.-M., Lu, N.T., Knapp, R., Chantanaorrapint, S., Limpanasittichai, P., Sun, H. , Gao, X.-F. & Zhang, L.-B. (2020). Evolutionary relationships of the ancient fern lineage the adder's tongues (Ophioglossaceae) with description of Sahashia gen. nov. Cladistics 36: 380–393. Https://doi.org/10.1111/cla.12408
7.  Zhang, L., Zhou, X.-M., Liang, Z.-L., Fan, X.-P., Lu, N.T., Song, M.-S., Knapp, R., Gao, X.-F. , Sun, H. & Zhang, L.-B. (2020). Phylogeny and classification of the tribe Lepisoreae (Polypodiaceae; pteridophyta) with the description of a new genus, Ellipinema gen. nov., segregated from Lepisorus. Molecular Phylogenetics and Evolution 148: 106803. https://doi.org/10.1016/j.ympev.2020.106803
8. Zhang, L., Lu, N.T., Zhou, X.-M., Chen, D.-K., Knapp, R., Zhou, L., Guo, L., Luong, T.T., Sun, H., Gao, X.-F. & Zhang, L.-B. (2019). A plastid phylogeny of the Old World fern genus Leptochilus (Polypodiaceae): Implications for cryptic speciation and progressive colonization from lower to higher latitudes. Molecular Phylogenetics and Evolution 134: 311–322. https://doi.org/10.1016/j.ympev.2019.01.013
9. Zhang, L. & Zhang, L.-B. (2018). A classification of the fern genus Tectaria (Tectariaceae; Polypodiales). Annals of the Missouri Botanical Garden 103: 188–199. https://doi.org/10.3417/2017007
10. Zhou, X.-M., Zhang, L., Lu, N.T., Gao, X.-F. & Zhang, L.-B. (2018). Pteridryaceae—A new fern family of Polypodiineae (Polypodiales) including taxonomic treatments. Journal of Systematics and Evolution 56: 148–173. https://doi.org/10.1111/jse.12305
11. Zhang, L. & Zhang, L.-B. (2018). Phylogeny and systematics of the brake fern genus Pteris (Pteridaceae) based on molecular (plastid and nuclear) and morphological evidence. Molecular Phylogenetics and Evolution 118: 265–285. http://dx.doi.org/10.1016/j.ympev.2017.09.011
12. Zhang, L., Zhou, X.-M., Chen, D.-K., Schuettpelz, E., Knapp, R., Lu, N.T., Luong, T.T., Dang, M.T., Duan, Y.-F., He, H., Gao, X.-F. & Zhang, L.-B. (2017). A global phylogeny of the fern genus Tectaria (Tectariaceae: Polypodiales) based on plastid and nuclear markers identifies major evolutionary lineages and suggests repeated evolution of free venation from anastomosing venation. Molecular Phylogenetics and Evolution 114: 295–333. http://dx.doi.org/10.1016/j.ympev.2017.05.020
13. Zhang, L., Zhou, X.-M., Lu, N.T. & Zhang, L.-B. (2017). Phylogeny of the fern subfamily Pteridoideae (Pteridaceae; Pteridophyta), with the description of a new genus: Gastoniella. Molecular Phylogenetics and Evolution 109: 59–72. http://dx.doi.org/10.1016/j.ympev.2016.12.037
14. Zhang, L., Schuettpelz, E., Rothfels, C. J., Zhou, X.-M., Gao, X.-F. & Zhang, L.-B. (2016). Circumscription and phylogeny of the fern family Tectariaceae based on plastid and nuclear markers, with the description of two new genera: Draconopteris and Malaifilix (Tectariaceae). Taxon 65: 723–738. https://doi.org/10.12705/654.3
15. Zhang, L., Rothfels, C. J., Ebihara, A., Schuettpelz, E., Péchon, T. L., Kamau, P., He, H., Zhou, X.-M., Prado, J., Field, A., Yatskievych, G., Gao, X.-F. & Zhang, L.-B. (2015). A global plastid phylogeny of the brake fern genus Pteris (Pteridaceae) and related genera in the Pteridoideae. Cladistics 31: 406–423. https://doi.org/10.1111/cla.12094
16. Zhang, L.-B. & Zhang, L. (2015). Didymochlaenaceae: A new fern family of eupolypods I (Polypodiales). Taxon 64: 27–38. http://dx.doi.org/10.12705/641.4
17.  Zhang, L.-B. & Zhang, L. (2012). The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris: The molecular phylogeny, systematics, and nomenclature of Dryopteris subg. Nothoperanema (Dryopteridaceae). Taxon 61: 1199–1216. https://doi.org/10.1002/tax.616003

• ZHANG Liang, YU Zhiyong, LU Thi Ngan. 2026. Pteridophytes of Xilong Mountain (西隆山蕨类植物Các loài dương xỉ và quyết thực vật ở núi Xilong). Kunming: Yunnan Fine Arts Publishing House. (In Press)

• 李恒, 李嵘，马文章，张良，王欣宇. 2020. 高黎贡山植物资源与区系地理. 武汉：湖北科学技术出版社.

• 税玉民，陈文红，武建勇，司马永康，杨治国，张良，马文章等. 2020. 云南河口县野生高等植物及植被多样性. 昆明：云南科技出版社.

目前正聚焦于喜马拉雅边境地区、滇东南、老挝和越南等区域的蕨类植物多样性调查，以及薄唇蕨属等类群的系统分类与进化研究。

现指导学生：

张鹭璐，硕士研究生，中国科学院大学

靳双宇，硕士研究生，云南大学

陆祖茂，硕士研究生，云南大学

谭越友，本科生，普洱学院

协助完成指导学生：

·2015–19: 周新茂，博士，中国科学院成都生物研究所；卷柏属的系统分类研究

·2016–20: Ngan Thi Lu，博士，中国科学院成都生物研究所/越南自然历史博物馆；复叶耳蕨属的系统分类研究

·2019–21: 范雪萍，硕士，中国科学院成都生物研究所；肿足蕨科的系统分类研究

·2020–23: 万霞，博士，中国科学院成都生物研究所/密苏里植物园；叉蕨科的生物地理学研究

·2020–23: 邱永玲，硕士，云南大学；西藏墨脱鳞毛蕨科和蹄盖蕨科的多样性研究

·2020–23: 杨建军，硕士，云南大学；禾叶蕨类的系统分类研究